Carrying energy data he has collected in space, Cosmos desperately attempts to keep it safe as Astrotrain, Starscream, and Thrust give chase. Warned not to transfer the data to Autobot Headquarters as long as the Decepticons are within range, Cosmos is shot down and crashes on a world known as Titan. Impoverished and barren, the planet is home to poverty-stricken natives who suffer under the rule of astrological priests and the enforced sacrifice of what little harvest they accrue to idols created in the image of so-called sky gods. The unconscious Cosmos is taken for such a god and duly revered, but not by all. A forceful female warrior named Talaria refuses to believe in the preachings of Jerro, one of the priests, and accuses him of using the faith of the natives for his own personal gain. As this happens, the Decepticons land, and Astrotrain quickly becomes a pretender to the heavenly rule, gaining Jerro's unctuous servitude and the terrified obedience of the people. Jerro takes Astrotrain to an incalculable amount of unstable energy crystals, and realizing that they are the source of Cosmos's data, the Decepticon orders the natives to stockpile them. Meanwhile, Talaria interacts with the hapless Cosmos and sends a homing signal to Earth, which results in Omega Supreme making an energy-draining flight to Titan with Jazz and Perceptor aboard. The Decepticons later take Talaria prisoner, but even as Omega Supreme sends Astrotrain's group clambering for the firmament, Jazz saves her before she can be sacrificed to strike further fear into the populace. A final blast from Astrotrain detonates the unstable crystals, making the natives' land uninhabitable. However, the Autobots take them to a safer part of the planet, giving them a new home and a new outlook on life.